Ownership Submission
FORM 4
Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).
UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940
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(Print or Type Responses)
1. Name and Address of Reporting Person *
WHITE TONY L
   2. Issuer Name and Ticker or Trading Symbol
APPLERA CORP [ABI/CRA] 		5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)
__X__ Director _____ 10% Owner
__X__ Officer (give title below) _____ Other (specify below)
Chairman, President and CEO
(Last)
(First)
(Middle)
APPLERA CORPORATION, 301 MERRITT 7		 3. Date of Earliest Transaction (Month/Day/Year)
05/02/2007
(Street)

NORWALK, CT 06851-1070		 4. If Amendment, Date Original Filed(Month/Day/Year)
 6. Individual or Joint/Group Filing(Check Applicable Line)
_X_ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person
(City)
(State)
(Zip)
Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1.Title of Security
(Instr. 3)		 2. Transaction Date (Month/Day/Year) 2A. Deemed Execution Date, if any (Month/Day/Year) 3. Transaction Code
(Instr. 8)		 4. Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)		 5. Amount of Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 3 and 4)		 6. Ownership Form: Direct (D) or Indirect (I)
(Instr. 4)		 7. Nature of Indirect Beneficial Ownership
(Instr. 4)
Code V Amount (A) or (D) Price
Applied Biosystems Group Common Stock 05/02/2007   M   46,991 A $ 17.98 476,657.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   939 D $ 31.01 475,718.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   235 D $ 31 475,483.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   704 D $ 30.99 474,779.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   1,408 D $ 30.96 473,371.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   704 D $ 30.95 472,667.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   704 D $ 30.94 471,963.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   469 D $ 30.92 471,494.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   704 D $ 30.9 470,790.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   704 D $ 30.89 470,086.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   59 D $ 30.88 470,027.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   997 D $ 30.87 469,030.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   1,936 D $ 30.86 467,094.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   3,813 D $ 30.85 463,281.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   4,752 D $ 30.84 458,529.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   3,284 D $ 30.83 455,245.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   2,757 D $ 30.82 452,488.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   4,576 D $ 30.81 447,912.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   7,218 D $ 30.8 440,694.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   4,341 D $ 30.79 436,353.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   3,930 D $ 30.78 432,423.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   1,877 D $ 30.77 430,546.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   411 D $ 30.76 430,135.3343 D  
Applied Biosystems Group Common Stock 05/02/2007   S(1)   469 D $ 30.75 429,666.3343 D  
Applied Biosystems Group Common Stock               75,000 I By Elizabeth Ann White Business Interests, LLLP (2)
Applied Biosystems Group Common Stock               63,000 I By Tony Lee White 2006 Two Year Grantor Retained Annuity Trust (3)
Applied Biosystems Group Common Stock               267,000 I By Tony Lee White 2007 Grantor Retained Annuity Trust (4)
Celera Group Common Stock 05/02/2007   M   11,748 A $ 7.3301 106,795.0998 D  
Celera Group Common Stock 05/02/2007   S(1)   715 D $ 14.37 106,080.0998 D  
Celera Group Common Stock 05/02/2007   S(1)   774 D $ 14.36 105,306.0998 D  

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Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned
(e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security
(Instr. 3)		 2. Conversion or Exercise Price of Derivative Security 3. Transaction Date (Month/Day/Year) 3A. Deemed Execution Date, if any (Month/Day/Year) 4. Transaction Code
(Instr. 8)		 5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4, and 5)		 6. Date Exercisable and Expiration Date
(Month/Day/Year)		 7. Title and Amount of Underlying Securities
(Instr. 3 and 4)		 8. Price of Derivative Security
(Instr. 5)		 9. Number of Derivative Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 4)		 10. Ownership Form of Derivative Security: Direct (D) or Indirect (I)
(Instr. 4)		 11. Nature of Indirect Beneficial Ownership
(Instr. 4)
Code V (A) (D) Date Exercisable Expiration Date Title Amount or Number of Shares
Applied Biosystems Group Employee Stock Options-Right to Buy $ 17.98 05/02/2007   M     46,991 06/26/2000 06/26/2007 Applied Biosystems Group Common Stock 46,991 $ 0 20,974 D  
Celera Group Employee Stock Options-Right to Buy $ 7.3301 05/02/2007   M     11,748 06/26/2000 06/26/2007 Celera Group Common Stock 11,748 $ 0 5,244 D  

Reporting Owners

Reporting Owner Name / Address Relationships
 Director  10% Owner  Officer  Other
WHITE TONY L
APPLERA CORPORATION
301 MERRITT 7
NORWALK, CT 06851-1070		   X     Chairman, President and CEO  

Signatures

 /s/ Thomas P. Livingston, Attorney-in-Fact for Tony L. White   05/04/2007
**Signature of Reporting Person Date

Explanation of Responses:

* If the form is filed by more than one reporting person, see Instruction 4(b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
(1) Reflects sale by independent third-party administrator under the issuer's Insider Diversification Program.
(2) Elizabeth Ann White Business Interests, LLLP, is a family limited parnership (the "FLP"). The reporting person's adult daughter is the sole general partner of the FLP, and the reporting person's wife and a grantor retained annuity trust established by the reporting person are currently the sole limited partners of the FLP.
(3) The reporting person's wife is the sole trustee of the Tony Lee White 2006 Two Year Grantor Retained Annuity Trust.
(4) The reporting person's wife is the sole trustee of the Tony Lee White 2007 Grantor Retained Annuity Trust.
 
Remarks:
This is the first of two forms being filed by the reporting person on May 4, 2007.

Note: File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.

Potential persons who are to respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB number.